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Gnus will read the mail spool when you activate a mail group. The mail file is first copied to your home directory. What happens after that depends on what format you want to store your mail in.
There are six different mail back ends in the standard Gnus, and more
back ends are available separately. The mail back end most people use
(because it is possibly the fastest) is nnml
(see section 6.3.13.3 Mail Spool).
6.3.13.1 Unix Mail Box Using the (quite) standard Un*x mbox. 6.3.13.2 Rmail Babyl Emacs programs use the Rmail Babyl format. 6.3.13.3 Mail Spool Store your mail in a private spool? 6.3.13.4 MH Spool An mhspool-like back end. 6.3.13.5 Maildir Another one-file-per-message format. 6.3.13.10 Mail Folders Having one file for each group. 6.3.13.11 Comparing Mail Back Ends An in-depth looks at pros and cons.